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Class Graptolithoidea, Order Rhabdopleuroidea, Family Uncertain

                        
Rhabdotubus johanssoni Bengtson & Urbanek, 1986
Fragments of erect zooidal tubes. of Rhabdotubus johanssoni.
Above, primary adhesion and later divergence (dv) of two adjacent tubus (t). To the right is a fragment of the tube in transmitted light.
Rhabdotubus johanssoni is regarded as the earliest known pterobranch. It was described from the early Middle Cambrian Eccaparadoxides pinus Zone of Närke, southern Sweden, by Stefan Bengtson and Adam Urbanek, and named in honour of its discoverer, Mr. Jan Johansson of Sköllersta, Sweden.
    
Bengtson & Urbanek (1986:293) wrote:

"The colonies encrusted shells of inarticulate brachiopods, and occasionally trilobites, on otherwise soft substrates. The tubarium consists  of repent and erect tubes. The former branch irregularly and produce a thecorhiza-like structure; the latter are erect and mostly isolated, up to 10 mm long in length and widening gradually to about 1 mm width. Both erect and repent tubes are composed of fusellar bands, mostly irregularly arranged. Branching of repent tubes takes place through resorption or perforation of fusellar tissue in the parent tube. Branching of erect tubes occurs sporadically. There is no thecal dimorphism. No sclerotized stolon is present."
The earliest known pterobranch hemichordate ! A number of similarities between this rhabdopleurid and the Tuboidea, an order of sessile graptolites, may indicate a close phylogenetic relationship between Rhabdotubus and tuboid graptolites.
BIBLIOGRAPHY

Bengtson, A. & Urbanek, A. 1986.
Rhabdotubus, a Middle Cambrian rhabdopleurid hemichordate. - Lethaia 19, 293-308.

Urbanek, A., Mierzejewski, P. & Bengtson, S. 1992. Fine structure of the earliest known rhabdopleurid. -
Lethaia, 25, 349-350.
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2003-2006
Graptolite Net - Portal of Graptolites and Pterobranchs.
Taxonomy of the order Rhabdopleuroidea (= Rhabdopleurida).
Taxonomy of the order Cephalodiscoidea (= Cephalodiscida).
Rhabdopleura kozlowskii Kulicki, 1969 - the earliest known member of the genus Rhabdopleura.
Cephalodiscus lutetianus Abrard et al. 1950 - the earliest known member of the genus Cephalodiscus ?
Diplohydra szaniawskii Mierzejewski et Kulicki, 2002 - a Permian hemichordate.
Graptolite-like pattern in the fusellar tissue of Paleozoic rhabdopleurid pterobranchs.
Pterobranchites antiquus Kozlowski, 1967 - an Ordovician Cephalodiscus-like hemichordate.
SEM micrographs of Rhabdopleura compacta
Rhabdopleuroididae - Ordovician rhabdopleurid pterobranchs.
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